2020 in paleobotany

From Wikipedia the free encyclopedia

List of years in paleobotany
In paleontology
2017
2018
2019
2020
2021
2022
2023
In arthropod paleontology
2017
2018
2019
2020
2021
2022
2023
In paleoentomology
2017
2018
2019
2020
2021
2022
2023
In paleomalacology
2017
2018
2019
2020
2021
2022
2023
In reptile paleontology
2017
2018
2019
2020
2021
2022
2023
In archosaur paleontology
2017
2018
2019
2020
2021
2022
2023
In mammal paleontology
2017
2018
2019
2020
2021
2022
2023
In paleoichthyology
2017
2018
2019
2020
2021
2022
2023

This article records new taxa of fossil plants that are scheduled to be described during the year 2020, as well as other significant discoveries and events related to paleobotany that are scheduled to occur in the year 2020.

Flowering plants[edit]

Alismatales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Limnobiophyllum pedunculatum[1]

Sp. nov

Valid

Low, Su & Xing in Low et al.

Late Oligocene

 China

A member of the family Araceae.

Apiales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Paleopanax puryearensis[2]

Sp. nov

Valid

Na, Blanchard & Wang

Middle Eocene

Cockfield Formation

 United States
( Tennessee)

A member of the family Araliaceae.

Arecales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Echimonocolpites chicxulubensis[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of a flowering plant, probably a member of the family Arecaceae.

Palmoxylon ceroxyloides[4]

Sp. nov

In press

Khan, Hazra & Bera in Khan et al.

Late Cretaceous (Maastrichtian)-Paleocene (Danian)

Deccan Intertrappean Beds

 India

A petrified palm stem of a member of the subfamily Ceroxyloideae.

Palmoxylon dindoriensis[5]

Sp. nov

Valid

Khan, Roy & Bera in Khan et al.

Late Cretaceous (Maastrichtian)-Paleocene (Danian)

Deccan Intertrappean Beds

 India

A petrified palm stem.

Sabalites dawsonii[6]

Sp. nov

Valid

Greenwood & Conran

Eocene

 Canada
( British Columbia)

Sabalites karondiensis[7]

Sp. nov

In press

Roy, Hazra & Khan in Roy et al.

Late Cretaceous-Paleocene (latest Maastrichtian-earliest Danian)

Deccan Intertrappean Beds

 India

A palm frond .

Spinizonocolpites riochiquensis[8]

Sp. nov

Valid

Vallati & De Sosa Tomas in Vallati, De Sosa Tomas & Casal

Late Cretaceous (Maastrichtian)

Lago Colhué Huapí Formation

 Argentina

A member of Arecaceae described on the basis of fossil pollen grains. Announced in 2019; the final version of the article naming it was published in 2020.

Buxales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Pachysandra europaea[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Pachysandra. Announced in 2019; the final version of the article naming it was published in 2020.

Caryophyllales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Gomphrenipollis garciae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant, possibly produced by members of the family Amaranthaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Chloranthales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Sarcandraxylon[11]

Gen. et sp. nov

Valid

Pipo, Iglesias & Bodnar

Late Cretaceous (early–middle Campanian)

Santa Marta Formation

Antarctica

A member of the family Chloranthaceae. Genus includes new species S. sanjosense.

Cornales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Amersinia littletonensis[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

Langtonia parva[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

A member of the family Mastixiaceae.

Mastixicarpum hoodii[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

A member of the family Mastixiaceae.

Nyssa gergoei[13]

Sp. nov

Valid

Hably

Miocene

 Hungary

A tupelo.

Nyssa gyoergyi[13]

Sp. nov

Valid

Hably

Miocene

 Hungary

A tupelo.

Platycrater iljinskajae[14]

Sp. nov

In press

Denk et al.

Late Oligocene

 Russia

Portnallia alexanderi[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

A member of the family Mastixiaceae.

Crossosomatales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images
Staphylea woodworthensis[15] Sp. nov In press Zhu & Manchester Oligocene Renova Formation  United States
( Montana)
A species of Staphylea.

Cucurbitales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Coriaripites goodii[16]

Sp. nov

Valid

Barreda, Palazzesi & Tellería in Renner et al.

Late Cretaceous (Campanian–early Maastrichtian)

Lopez de Bertodano Formation
Santa Marta Formation
Snow Hill Island Formation

Antarctica

Pollen grains similar to those of extant members of the genus Coriaria.

Echitriporites jolyi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Cayaponia. Announced in 2020; the final version of the article naming it was published in 2021.

Ericales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Andrewsiocarpon puryearensis[2]

Sp. nov

Valid

Na, Blanchard & Wang

Middle Eocene

Cockfield Formation

 United States
( Tennessee)

A member of the family Theaceae.

Anubiscarpon[17]

Gen. et sp. nov

Valid

Smith & Manchester

Middle Eocene

Clarno Formation

 United States
( Oregon)

A member of the family Theaceae. Genus includes new species A. andersonae.

Fabales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Cercis zekuensis[18]

Sp. nov

Valid

Li et al.

Early Miocene

 China

A species of Cercis. Announced in 2020; the final version of the article naming it was published in 2021.

Gleditsia pliocaenica[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Gleditsia. Announced in 2019; the final version of the article naming it was published in 2020.

Menendoxylon lutzi[19] Sp. nov In press Baez & Crisafulli Miocene Chiquimil  Argentina Fossil wood of a member of the family Fabaceae.

Parkiidites marileae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Parkia. Announced in 2020; the final version of the article naming it was published in 2021.

Prioria martineziorum[20]

Sp. nov

Valid

Rodríguez-Reyes & Estrada-Ruiz

Oligocene-Miocene

Santiago Formation

 Panama

A species of Prioria.

Psilastephanocolporites deoliverae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the family Polygalaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Psilastephanocolporites endoporatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Striatopollis grahamii[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of an eudicot, probably a member of the family Fabaceae.

Fagales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Alnus chaybulakensis[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

 Kazakhstan

An alder.

Berryophyllum hainanensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A member of the family Fagaceae.

Carpinus asymmetrica[23]

Sp. nov

In press

Xue & Jia in Xue et al.

Early Miocene

Maguan

 China

A species of Carpinus.

Carpinus symmetrica[23]

Sp. nov

In press

Xue & Jia in Xue et al.

Early Miocene

Maguan Basin

 China

A species of Carpinus

Carya pipecreekensis[24]

Sp. nov

In press

Swinehart & Farlow

Late Neogene

Pipe Creek Sinkhole

 United States
( Indiana)

A hickory.

Castaneophyllum hainanensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A member of the family Fagaceae.

Castaneophyllum lanceolata[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A member of the family Fagaceae.

Castanopsis bulgarica[25] Sp. nov Valid Mantzouka, Ivanov & Bozukov Late Miocene–early Pliocene (late Messinian–early Zanclean) Pokrovnik  Bulgaria A species of Castanopsis. Announced in 2020; the final version of the article naming it was published in 2021.

Lithocarpus changchangensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A species of Lithocarpus.

Quercus borissovii[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

 Kazakhstan

An oak.

Quercus changchangensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

Quercus paleoargyrotricha[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

Quercus paleohypargyrea[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

Quercus paleolamellosa[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

Garryales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Eucommia szaferi[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Eucommia. Announced in 2019; the final version of the article naming it was published in 2020.

Gentianales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Asclepiadospermum[26]

Gen. et 2 sp. nov

Valid

Del Rio et al.

Early Eocene

Niubao Formation

 China

An asclepiadoid Apocynaceae genus.
Included species A. marginatum and A. ellipticum.

Margocolporites carinae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Rauvolfia. Announced in 2020; the final version of the article naming it was published in 2021.

Icacinales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Iodes elliptica[27]

Sp. nov

In press

Del Rio et al.

Early Oligocene

Wenshan Basin

 China

A member of the family Icacinaceae.

Iodes passiciensis[28]

Sp. nov

Valid

Del Rio & De Franceschi

Early Eocene

 France

A member of the family Icacinaceae.

Manchesteria[29]

Gen. et sp. nov

In press

Stull & Rozefelds in Rozefelds et al.

Cenozoic (mid-Miocene or, more likely, middle Eocene)

 Australia

A member of the family Icacinaceae. Genus includes new species M. australis.

Pyrenacantha simonsii[30]

Sp. nov

Valid

Stull et al.

Early Oligocene

 Egypt

A species of Pyrenacantha.

Lamiales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Echitricolpites cruziae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Aegiphila. Announced in 2020; the final version of the article naming it was published in 2021.

Multiareolites? reticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the family Acanthaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Retistephanocolpites curvimuratus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Retistephanocolpites pardoi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant, possibly produced by members of the genus Amphilophium. Announced in 2020; the final version of the article naming it was published in 2021.

Laurales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Actinodaphnoxylon[31]

Gen. et sp. nov

In press

Akkemik et al.

Eocene (Lutetian)

 Turkey

A member of the family Lauraceae. Genus includes new species A. zileensis.

Mezilaurinoxylon oleiferum[32]

Sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

 Argentina

A member of the family Lauraceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Patagonoxylon[32]

Gen. et sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

 Argentina

A member of Laurales of uncertain phylogenetic placement. Genus includes new species P. scalariforme. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Thymolepis[33]

Gen. et sp. nov

Valid

Chambers & Poinar

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Possibly an early representative of Monimiaceae. Genus includes new species T. toxandra.

Valviloculus[34]

Gen. et sp. nov

Valid

Poinar et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Possibly a member of Laurales related to the families Monimiaceae and Atherospermataceae. Genus includes new species V. pleristaminis.

Liliales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Smilax fujianensis[35]

Sp. nov

Valid

Dong et al.

Middle Miocene

 China

A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.

Smilax zhangpuensis[35]

Sp. nov

Valid

Dong et al.

Middle Miocene

 China

A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.

Magnoliales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Magnolia nanningensis[36]

Sp. nov

Valid

Huang et al.

Late Oligocene

Nanning Basin

 China

A species of Magnolia.

Magnolia waltheri[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Magnolia. Announced in 2019; the final version of the article naming it was published in 2020.

Melloniflora[37]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

Potomac Group

 United States
( Virginia)

A relative of extant early-diverging members of the Magnoliales. Genus includes new species M. virginiensis.

Malpighiales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Salix palaeofutura[38]

Sp. nov

Valid

Narita et al.

Miocene

Bifuka Formation

 Japan

A willow.

Malvales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images
Bastardioxylon[19] Gen. et sp. nov In press Baez & Crisafulli Miocene Chiquimil  Argentina Fossil wood of a member of the family Malvaceae. Genus includes new species B. antiqua.
Dipterocarpus dindoriensis[39] Sp. nov Valid Khan, Spicer & Bera in Khan et al. Late Cretaceous (Maastrichtian) Deccan Intertrappean Beds  India A species of Dipterocarpus.

Echiperiporites germeraadii[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Echiperiporites jaramilloi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Hibiscus. Announced in 2020; the final version of the article naming it was published in 2021.

Echiperiporites titanicus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Malachra. Announced in 2020; the final version of the article naming it was published in 2021.

Retistephanocolporites elizabeteae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Ceiba. Announced in 2020; the final version of the article naming it was published in 2021.

Veraguasoxylon[20]

Gen. et sp. nov

Valid

Rodríguez-Reyes & Estrada-Ruiz

Oligocene-Miocene

Santiago Formation

 Panama

A member of the family Malvaceae. Genus includes new species V. panamense.

Myrtales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Eucalyptus xoshemium[40]

Sp. nov

Valid

Gandolfo & Zamaloa in Zamaloa, Gandolfo & Nixon

Eocene (Ypresian)

Huitrera Formation

 Argentina

A species of Eucalyptus.

Mangroveoxylon[41]

Gen. et comb. nov

In press

Moya & Brea

Late Miocene?

Ituzaingó Formation

 Argentina

A member of the family Combretaceae; a new genus for "Menendoxylon" areniensis Lutz (1979).

Myrceugenellites grandiporosum[32]

Sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

 Argentina

A member of the family Myrtaceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Primotrapa[42]

Gen. et sp. nov

Valid

Li & Li in Li et al.

Late Eocene to early Miocene

Hannuoba

 China
 Czech Republic
 France
 Germany

A member of Trapoideae. Genus includes new species P. weichangensis, as well as "Carpolithus" pomelii Saporta (1878) and "Hemitrapa" alpina Su & Zhou in Su et al. (2018).

Nymphaeales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Praenymphaeapollenites[43]

Gen. et sp. nov

Valid

Barrón, Peris & Labandeira in Peris et al.

Cenomanian

Burmese amber

 Myanmar

Pollen of a member of Nymphaeaceae.
Genus includes new species P. cenomaniensis.

Oxalidales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Cunoniantha[44]

Gen. et sp. nov

Valid

Jud & Gandolfo

Paleocene (early Danian)

Salamanca Formation

 Argentina

A member of the family Cunoniaceae. Genus includes new species C. bicarpellata. Announced in 2020; the final version of the article naming it was published in 2021.

Elaeocarpus nanningensis[45]

Sp. nov

In press

Liu et al.

Late Oligocene

Yongning Formation

 China

A species of Elaeocarpus.

Elaeocarpus prelacunosus[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

Elaeocarpus preprunifolioides[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

Elaeocarpus prerugosus[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

Elaeocarpus preserratus[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

Elaeocarpus presikkimensis[45]

Sp. nov

In press

Liu et al.

Miocene

Erzitang Formation

 China

A species of Elaeocarpus.

Poales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Monoporopollenites scabratus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the family Poaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Rhizomatites[46]

Gen. et sp. nov

In press

Robledo & Anzótegui in Robledo et al.

Miocene-Pliocene

 Argentina

A member of Cyperaceae. Genus includes new species R. cyperoides.

Proteales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Banksia microphylla[47]

Sp. nov

Valid

Carpenter in Carpenter & Milne

Late Eocene

 Australia

A species of Banksia.

Banksieaeidites zanthus[47]

Sp. nov

Valid

Milne in Carpenter & Milne

Late Eocene

 Australia

A Banksia-like pollen.

Platanus emryi[48]

Sp. nov

Valid

Huegele, Spielbauer & Manchester

Miocene

 United States

A species of Platanus.

Ranunculales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Cissampelos defranceschii[49]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A species of Cissampelos. Announced in 2020; the final version of the article naming it was published in 2021.

Clematis csabae[13]

Sp. nov

Valid

Hably

Miocene

 Hungary

A species of Clematis.

Diploclisia praeaffinis[50]

Sp. nov

Valid

Jia et al.

Late Miocene

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Menispermites bangorensis[49]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Menispermites haominae[49]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Menispermites tibetica[49]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Paleoorbicarpum[51]

Gen. et sp. nov

Valid

Han et al.

Paleocene

Sanshui Basin

 China

A member of the family Menispermaceae. Genus includes new species P. parvum.

Stephania bangorensis[49]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.

Stephania geniculata[51]

Sp. nov

Valid

Han et al.

Paleocene

Sanshui Basin

 China

A species of Stephania

Stephania ornamenta[51]

Sp. nov

Valid

Han et al.

Paleocene

Sanshui Basin

 China

A species of Stephania

Stephania shuangxingii[49]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.

Rosales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Berhamniphyllum junrongii[52]

Sp. nov

Valid

Zhou, Wang & Huang in Zhou et al.

Late Eocene

Markam Basin

 China

A member of the family Rhamnaceae

Crataegus pentagynoides[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Crataegus. Announced in 2019; the final version of the article naming it was published in 2020.

Hemiptelea kryshtofovichii[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

 Kazakhstan

A member of the family Ulmaceae.

Scabrastephanoporites[3]

Gen. et sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of an eudicot, probably a member of the family Ulmaceae or Cannabaceae. Genus includes new species S. variabilis.

Sapindales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Acer dombeyopsis[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

Acer viburnoides[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

Acer vitiforme[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

Brosipollis reticulatus[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of a flowering plant, probably a member of the family Burseraceae.

Choerospondias fujianensis[53]

Sp. nov

In press

Wang et al.

Miocene

 China

A species of Choerospondias.

Llanodelacruzoxylon[54] Gen. et sp. nov Rodríguez-Reyes, Estrada-Ruiz & Gasson Oligocene–Miocene Santiago Formation  Panama A member of the family Anacardiaceae. Genus includes new species L. sandovalii.

Manchestercarpa[55]

Gen. et sp. nov

Valid

Atkinson

Late Cretaceous (Campanian)

 Canada
( British Columbia)

A member of the family Meliaceae described on the basis of a fossil fruit. Genus includes new species M. vancouverensis.

Parametopioxylon[56]

Gen. et sp. nov

Valid

Franco et al.

Miocene

Ituzaingó Formation

 Argentina

A member of the family Anacardiaceae described on the basis of fossil wood. Genus includes new species P. crystalliferum.

Psilastephanocolporites hammenii[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of a flowering plant, probably a member of the family Meliaceae

Quinquala[57]

Gen. et sp. nov

Valid

Manchester & Disney in Manchester, Disney & Pham

Eocene

Clarno Formation
Tepee Trail Formation

 United States
( Oregon
 Wyoming)

A fossil fruit with affinities with the Rutaceae. Genus includes new species Q. obovata.

Rousea cavitata[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Saxifragales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Corylopsis grisea[58]

Sp. nov

Valid

Quirk & Hermsen

Early Pliocene

Gray Fossil Site

 United States
( Tennessee)

A species of Corylopsis. Announced in 2020; the final version of the article naming it was published in 2021.

Itea polyneura[59]

Sp. nov

In press

Huang & Tian in Tian et al.

Oligocene

Huazhige Formation

 China

A species of Itea.

Protoaltingia[60]

Gen. et sp. nov

Valid

Scharfstein, Stockey & Rothwell

Late Cretaceous (Coniacian)

 Canada
( British Columbia)

A member of the family Altingiaceae. Genus includes new species P. comoxense.

Solanales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Physalis hunickenii[61]

Sp. nov

Valid

Deanna, Wilf & Gandolfo

Early Eocene

Laguna del Hunco Formation

 Argentina

A species of Physalis.

Trochodendrales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Eotrochion[62]

Gen. et sp. nov

Valid

Manchester, Kvaček & Judd

Paleocene

 United States
( Wyoming)

A member of the family Trochodendraceae. Genus includes new species E. polystylum. Announced in 2020; the final version of the article naming it was published in 2021.

Paraconcavistylon[62]

Gen. et comb. nov

Valid

Manchester, Kvaček & Judd

Ypresian

Klondike Mountain Formation

 United States
( Washington)

A Trochodendraceae; a new genus for "Concavistylon" wehrii Manchester et al. (2018). Announced in 2020; the final version of the article naming it was published in 2021.

Trochodendron infernense[62]

Sp. nov

Valid

Manchester, Kvaček & Judd

Paleocene

 United States
( Wyoming)

A species of Trochodendron. Announced in 2020; the final version of the article naming it was published in 2021.

Vitales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Yua jiangxiensis[63]

Sp. nov

Valid

He & Wang

Miocene

Toupi Formation

 China

A species of Yua. Announced in 2020; the final version of the article naming it was published in 2021.

Other angiosperms[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Aextoxicoxylon kawasianus[64]

Sp. nov

Valid

Vera et al.

Late Cretaceous

Puntudo Chico Formation

 Argentina

A fossil dicot wood

Atlantocarpus[65]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

 Portugal
 United States
( Virginia)

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species A. virginiensis.

Carpolithes gergoei[13]

Sp. nov

Valid

Hably & Erdei in Hably

Miocene

 Hungary

A fossil fruit of a flowering plant of uncertain phylogenetic placement.

Catanthus[66]

Gen. et sp. nov

In press

Friis, Crane & Pedersen

Early Cretaceous

 Portugal

An early flowering plant. Genus includes new species C. dolichostemon.

Cavilignum[67]

Gen. et sp. nov

Valid

Siegert & Hermsen

Early Pliocene

Gray Fossil Site

 United States
( Tennessee)

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil endocarps. Genus includes new species C. pratchettii.

Chainandra[68]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Genus includes new species C. zeugostylus.

Cichoreacidites? flammulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Crotonoidaepollenites echinatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Cyathitepala[69]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Genus includes new species C. papillosa.

Dasykothon[70]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A flowering plant of uncertain phylogenetic placement, possibly a member of Laurales. Genus includes new species D. leptomiscus.

Dinganthus[71]

Gen. et sp. nov

Valid

Liu et al.

Miocene

Dominican amber

 Dominican Republic

A eudicot of uncertain phylogenetic placement. Genus includes new species D. pentamera.

Echistephanoporites annulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Eofructus[72]

Gen. et sp. nov

Valid

Han & Wang

Early Cretaceous

Yixian Formation

 China

An infructescence including a central axis and five fruits resembling Liaoningfructus. Genus includes new species E. liutiaogouensis.

Foveotricolporites crassus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Inaperturopollenites microechinatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Ladakhipollenites carmoi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Lambertiflora[65]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

 United States
( Virginia)

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species L. elegans.

Malvacipolloides diversus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Malvacipolloides echibaculatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Malvacipolloides romeroae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Menatanthus[73]

Gen. et sp. nov

Valid

Uhl, Paudayal & El Atfy in Uhl et al.

Paleocene (Thanetian)

 France

A eudicot of uncertain phylogenetic placement. Genus includes new species M mosbruggeri. Announced in 2020; the final version of the article naming it was published in 2021.

Mugideiriflora[65]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Aptian-early Albian)

Almargem Formation

 Portugal

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species M. portugallica.

Phantophlebia[74]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A flowering plant of uncertain phylogenetic placement, possibly related to myrsinoid members of the family Primulaceae. Genus includes new species P. dicycla.

Platanites willigeri[75]

Sp. nov

Valid

Halamski & Kvaček in Halamski et al.

Late Cretaceous (Santonian)

Czerna Formation

 Poland

Trifoliolate platanoid leaves.

Psilaperiporites delicatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Psilaperiporites lunaris[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Ranunculacidites reticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Rasenganus[76]

Gen. et sp. nov

Valid

Xing & Gu

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A possible epizoochorous fruit. Genus includes new species R. auricularus.

Retibrevitricolpites microreticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retibrevitricolporites costaporus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retibrevitricolporites? toigoae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retipollenites solimoensis[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retitriporites crassoreticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Rhoipites alfredii[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Singpuria[77]

Gen. et sp. nov

Valid

Ramteke, Manchester & Nagrale in Ramteke et al.

Late Cretaceous (Maastrichtian)

Deccan Intertrappean Beds

 India

A member of Pentapetalae of uncertain phylogenetic placement. Genus includes new species S. kapgatei.

Sinoherba[78]

Gen. et sp. nov

Valid

Liu & Wang in Liu, Chen & Wang

Early Cretaceous (BarremianAptian)

Yixian Formation

 China

An early monocot. Genus includes new species S. ningchengensis. Announced in 2020; the final version of the article naming it was published in 2021.

Varifructus[79]

Gen. et sp. nov

In press

Liu et al.

Early Cretaceous

Yixian Formation

 China

An early flowering plant. Genus includes new species V. lingyuanensis.

Wireroadia[80]

Gen. et sp. et comb. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian to Santonian)

 United States
( Alabama
 Massachusetts
 New Jersey
 New York)

A winged fruit of a eudicot of uncertain phylogenetic placement. Genus includes new species W. viccallii, as well as W. major (Hollick).

Pinales[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Agathis ledongensis[81]

Sp. nov

Valid

Oskolski et al.

Late Oligocene–early Miocene

Qiutangling Formation

 China

A species of Agathis

Agathoxylon cozzoi[82]

Sp. nov

Valid

Gnaedinger & Zavattieri

Late Triassic

Chihuido Formation

 Argentina

Araucaria cuneoi[83]

Sp. nov

Valid

Noll & Kunzmann

Middle Jurassic

 Argentina

A species of Araucaria.

Araucaria famii[84]

Sp. nov

Valid

Stockey & Rothwell

Late Cretaceous (Campanian)

 Canada
( British Columbia)

A species of Araucaria.

Araucaria fildesensis[85]

Sp. nov

Valid

Shi et al.

Eocene

Fossil Hill Formation

Antarctica
(King George Island)

A species of Araucaria

Araucaria huncoensis[86]

Sp. nov

Valid

Rossetto‐Harris in Rossetto‐Harris et al.

Early Eocene

Laguna del Hunco Formation

 Argentina

A species of Araucaria.

Araucaria stockeyana[83]

Sp. nov

Valid

Noll & Kunzmann

Middle Jurassic

 Argentina

A species of Araucaria.

Araucarites pachacuteci[87] Sp. nov

In press

Martínez in Martínez et al.

Early Cretaceous (BerriasianValanginian)

Huancané Formation

 Peru

A member of the family Araucariaceae

Brachyoxylon zhouii[88]

Sp. nov

Valid

Jiang et al.

Early Cretaceous

Guantou Formation

 China

A conifer wood. Announced in 2020; the final version of the article naming it was published in 2021.

Brachyphyllum sattlerae[89] Sp. nov Valid Batista et al. Early Cretaceous (Aptian) Crato Formation  Brazil A member of the family Pinidae.

Callialastrobus[90]

Gen. et sp. nov

In press

Kvaček & Mendes

Early Cretaceous (late Aptian–early Albian)

Lusitanian Basin

 Portugal

A pollen cone of a member of Araucariaceae. Genus includes new species C. sousai.

Cedrus anatolica[91]

Sp. nov

Valid

Akkemik

Early Miocene

Hançili Formation

 Turkey

A species of Cedrus. Announced in 2020; the final version of the article naming it was published in 2021.

Circoporoxylon tibetense[92]

Sp. nov

In press

Xia et al.

Middle Jurassic (Callovian)

Xiali Formation

 China

Possibly a member of the family Podocarpaceae.

Comoxostrobus[93]

Gen. et sp. nov

Valid

Stockey, Rothwell & Atkinson

Late Cretaceous (early Coniacian)

 Canada
( British Columbia)

A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species C. rossii.

Cupressinoxylon klimovii[94]

Nom. nov

Valid

Blokhina

Miocene

 Russia

A member of the family Cupressaceae; a replacement name for Cupressinoxylon biotoides Blokhina (1989).

Cupressinoxylon llantenesense[82]

Sp. nov

Valid

Gnaedinger & Zavattieri

Late Triassic

Llantenes Formation

 Argentina

Cupressinoxylon manuelii[95] Sp. nov In press Ríos-Santos, Cevallos-Ferriz & Pujana Late Cretaceous (Campanian-Maastrichtian) Cabullona Group  Mexico

Ductoagathoxylon wangii[96]

Sp. nov

In press

Gou & Feng in Gou et al.

Middle Jurassic

Xishanyao Formation

 China

A conifer stem.

Friisia[97]

Gen. et sp. nov

In press

Mendes & Kvaček

Early Cretaceous (late Aptian – early Albian)

Lusitanian Basin

 Portugal

An ovuliferous cone of a member of the family Podocarpaceae. Genus includes new species F. lusitanica.

Juniperoxylon acarcaea[98]

Sp. nov

In press

Akkemik

Early Miocene

Hançili Formation

 Turkey

A member of the family Cupressaceae.

Lesbosoxylon kemaliyensis[99]

Sp. nov

Valid

Akkemik & Mantzouka in Akkemik, Mantzouka & Kıran Yıldırım

Miocene

Divriği Formation

 Turkey

A member of the family Pinaceae.

Marskea cuspidata[100]

Sp. nov

Valid

Frolov & Mashchuk

Middle Jurassic

Prisayan Formation

 Russia

A member of the family Taxaceae.

Mukawastrobus[101]

Gen. et sp. nov

Valid

Stockey, Nishida & Rothwell

Late Cretaceous (late Campanian—early Maastrichtian)

 Japan

A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species M. satoi.

Piceoxylon yumeniense[102]

Sp. nov

Valid

Zhou, Peng, Deng, Zhang & Yang in Zhou et al.

Early Cretaceous

Xiagou Formation

 China

Fossil wood of a member of the family Pinaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Pinuxylon selmeierianum[103]

Sp. nov

Valid

Dolezych & Reinhardt

Paleogene

Eureka Sound Group

 Canada
( Nunavut)

A member of the family Pinaceae described on the basis of fossil wood

Protophyllocladoxylon chijinense[102]

Sp. nov

Valid

Zhou, Peng, Deng, Zhang & Yang in Zhou et al.

Early Cretaceous

Xiagou Formation

 China

Announced in 2020; the final version of the article naming it was published in 2021.

Tsuga asiatica[104]

Sp. nov

Valid

Wu & Zhou in Wu et al.

Late Paleogene

 China

A species of Tsuga. Announced in 2019; the final version of the article naming it was published in 2020.

Other seed plants[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Amyelon turpanense[105]

Sp. nov

In press

Shi, Yu & Sun

Permian (Lopingian)

Wutonggou

 China

A root of a member of Cordaitales

Androstrobus obovatus[106]

Sp. nov

In press

Bodnar et al.

Late Triassic

Potrerillos Formation

 Argentina

A member of Cycadales.

Araripestrobus[107]

Gen. et sp. nov

Valid

Seyfullah, Roberts, Schmidt & Kunzmann in Seyfullah et al.

Early Cretaceous (Aptian-Albian)

Crato

 Brazil

A seed plant belonging to the group Erdtmanithecales. Genus includes new species A. resinosus.

Archaeopetalanthus[108]

Gen. et sp. nov

Valid

Naugolnykh

Carboniferous

Listvjanskaya

 Russia

A member of Pinophyta belonging to the group Vojnovskyales. Genus includes new species A. progressus.

Battenispermum[109]

Gen. et sp. nov

In press

Mendes, Pedersen & Friis

Early Cretaceous

 Portugal

A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species B. hirsutum.

Carpolithus volantus[110]

Sp. nov

Valid

Gómez et al.

Early Cretaceous (Aptian)

La Cantera Formation

 Argentina

A fossil seed, possibly produced by a member of Gnetales.

Ductolobatopitys[111]

Gen. et sp. nov

Valid

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

 Brazil

A gymnosperm described on the basis of fossil wood. Genus includes new species D. mussae.

Filigigantopteris[112]

Gen. et 2 sp. nov

In press

Zhou et al.

Late Permian

Nayixiong Formation

 China

A gigantopterid. Genus includes new species F. asymmetrica and F. dahaia.

Ginkgo pediculata[113]

Sp. nov

In press

Deng, Yang & Zhou

Early Cretaceous

 China

A species of Ginkgo.

Jianchangia[114]

Gen. et sp. nov

Valid

Yang, Wang & Ferguson

Early Cretaceous (Aptian)

Jiufotang Formation

 China

A member of Ephedraceae. Genus includes new species J. verticillata.

Johniphyllum[115]

Gen. et sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

 United States
( Texas)

A member of Voltziales. Genus includes new species J. multinerve.

Jordaniopteris[116]

Gen. et comb. nov

Valid

Anderson in Anderson et al.

Permian (possibly Lopingian)

Um Irna Formation

 Jordan

A seed fern. A new genus for "Dicroidium" irnensis Abu Hamad et al. (2008); genus also includes "Dicroidium" jordanensis Abu Hamad et al. (2008), "Dicroidium" robustum Kerp & Vörding (2008) and "Dicroidium" bandelii Abu Hamad et al. (2017).

Kaokoxylon brasiliensis[111]

Sp. nov

Valid

Conceição, Neregato & Iannuzzi in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

 Brazil

A conifer described on the basis of fossil wood.

Nilssonia mirovanae[117]

Sp. nov

Valid

Čepičková & Kvaček

Late Cretaceous (Cenomanian)

Peruc-Korycany Formation

 Czech Republic

A cycad.

Novaiorquepitys[118]

Gen. et sp. nov

In press

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

 Brazil

A gymnosperm stem. Genus includes new species N. maranhensis Conceição, Neregato & Iannuzzi.

Ovalocarpus butmanii[119]

Sp. nov

Valid

Naugolnykh & Linkevich

Permian (Artinskian)

 Russia
( Sverdlovsk Oblast)

A member of Ginkgoales belonging to the family Cheirocladaceae.

Palaeocupressinoxylon[120]

Gen. et sp. nov

In press

Wan, Yang & Wang

Late Permian

Turpan–Hami Basin

 China

A silicified gymnospermous fossil wood. Genus includes new species P. uniseriale.

Phoenicopsis anadyrensis[121]

Sp. nov

Valid

Nosova in Zolina et al.

Late Cretaceous–Paleocene (MaastrichtianDanian)

Rarytkin

 Russia
( Chukotka Autonomous Okrug)

A member of Czekanowskiales

Pseudovoltzia sapflorensis[115]

Sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

 United States
( Texas)

A member of Voltziales

Pterostoma neehoffii[122]

Sp. nov

Valid

Conran et al.

Middle Miocene

 New Zealand

A cycad

Pteruchus frenguellii[123]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

 Jordan

A pollen organ of a seed fern

Pteruchus lepidus[123]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

 Jordan

A pollen organ of a seed fern

Samaropsis jinchangensis[124]

Sp. nov

Valid

Hua & Sun in Hua et al.

Early Permian

 China

A seed fossil. Announced in 2019; the final version of the article naming was published in 2020.

Umaltolepis involuta[125]

Sp. nov

In press

Nosova

Middle Jurassic

Angren Formation

 Uzbekistan

Umaltolepis sogdianica[125]

Sp. nov

In press

Nosova

Middle Jurassic

Angren Formation

 Uzbekistan

Umkomasia aequatorialis[123]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

 Jordan

A cupulate structure of a seed fern

Wantus[115]

Gen. et sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

 United States
( Texas)

A member of Voltziales. Genus includes new species W. acaulis.

Wudaeophyton[126]

Gen. et sp. nov

Valid

Pšenička et al.

Early Permian

Taiyuan Formation

 China

A small vine, most similar to pteridosperms from the group Callistophytales. Genus includes new species W. wangii.

Yangopteris[127]

Gen. et comb. nov

In press

Zhou et al.

Permian (Asselian)

 China

A seed fern; a new genus for "Alethopteris" ascendens Halle.

Yvyrapitys[118]

Gen. et sp. nov

In press

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

 Brazil

A gymnosperm stem. Genus includes new species Y. novaiorquensis Conceição, Neregato & Iannuzzi.

Other plants[edit]

Name Novelty Status Authors Age Type locality Location Notes Images

Annularia paisii[128]

Sp. nov

Valid

Correia et al.

Carboniferous (Gzhelian)

 Portugal

Blasiites huolinhensis[129]

Sp. nov

In press

Li et al.

Early Cretaceous

Huolinhe Formation

 China

A liverwort belonging to the family Blasiaceae.

Botryopteris multifolia[130]

Sp. nov

In press

He et al.

Permian (Lopingian)

Junlian Formation

 China

A fern

Birisia mandshurica[131]

Sp. nov

Valid

Golovneva, Grabovskiy & Zolina

Early Cretaceous (Albian)

Frentsevka Formation

 Russia
( Primorsky Krai)

A fern belonging to the family Dicksoniaceae.

Calamites cambrensis[132]

Sp. nov

Valid

Thomas

Carboniferous (Pennsylvanian)

 United Kingdom

Catenuporella[133]

Gen. et sp. nov

Valid

Zhang et al.

Late Ordovician

Ordos Basin

 China

A green alga belonging to the group Dasycladales. Genus includes new species C. gigantia.

Charaxis spicatus[134]

emend. nov.

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Maestrat Basin

 Spain

Thallus of Echinochara lazarii. Both taxa has been firstly found anatomically attached

Chlamydomonas hanublikanus[135]

Sp. nov

Vršanská & Hinkelman

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A species of Chlamydomonas

Circinites[46]

Gen. et sp. nov

In press

Robledo & Anzótegui in Robledo et al.

Miocene-Pliocene

 Argentina

A fern belonging to the family Pteridaceae. Genus includes new species C. pteridoides.

Clavatisporites cenomaniana[136]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

 Argentina

Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Clavator calcitrapus var. jiangluoensis[134]

comb. nov

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Maestrat Basin

 Spain

Clavatoracean species.

Collarecodium? nezpercae[137] Sp. nov Valid Bucur & Rigaud in Bucur et al. Late Triassic (Norian)  United States
( Idaho)
A green alga belonging to the group Bryopsidales and possibly to the family Udoteaceae.

Collarisporites minor[136]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

 Argentina

Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Coniopteris sandaolingensis[138]

Sp. nov

Valid

Yuan & Sun in Yuan et al.

Middle Jurassic

Xishanyao Formation

 China

Dimicheleodendron[139]

Gen. et comb. nov

Valid

Thomas & Cleal

Carboniferous

 United Kingdom

A lycophyte; a new genus for "Lepidodendron" hickii.

Drynaria diplosticha[140]

Sp. nov

Valid

Yu & Xie in Yu et al.

Late Miocene

Bangmai Formation

 China

A fern belonging to the family Polypodiaceae.

Echinochara lazarii[134]

comb. nov

Valid

Pérez-Cano, Bover-Arnal et MArtín-Closas in Pérez-Cano et al.

Barremian

Maestrat Basin

 Spain

A member of Clavatoraceans.

Equicalastrobus pusillus[141]

Sp. nov

Valid

Zhang & Yan in Zhang et al.

Late Triassic

Baojishan Basin

 China

A member of Equisetales. Announced in 2020; the final version of the article naming was published in 2021.

Equisetum yenbaiense[142]

Sp. nov

Valid

Aung et al.

Late Miocene

 Vietnam

A species of Equisetum

Equisetum yongpingense[142]

Sp. nov

Valid

Aung et al.

Late Pliocene

Sanying Formation

 Vietnam

A species of Equisetum

Filippoporella[143] Gen. et sp. nov In press Sokač & Grgasović Early Paleocene  Croatia A green alga belonging to the group Dasycladales. Genus includes new species F. barattoloi.

Frullania partita[144]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A liverwort, a species of Frullania

Frullania vanae[145]

Sp. nov

Valid

Mamontov et al.

Eocene

Rovno amber

 Ukraine

A liverwort, a species of Frullania

Gippslandites[146]

Gen. et sp. nov

Valid

McSweeney, Shimeta & Buckeridge

Late Silurian–early Devonian

 Australia

A member of Zosterophyllaceae. Genus includes new species G. minutus.

Gmujij[147] Gen. et sp. nov Valid Pfeiler & Tomescu Devonian (Emsian) Battery Point  Canada
( Quebec)
An early euphyllophyte. Genus includes new species G. tetraxylopteroides. Announced in 2020; the final version of the article naming it was published in 2021.
Griphoporella minuta[137] Sp. nov Valid Bucur & Peybernes in Bucur et al. Late Triassic  Japan A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Hansopteris[148] Gen. et sp. nov In press Zhou et al. Permian (Asselian)  China An anachoropterid fern. Genus includes new species H. uncinatus.
Holosporella magna[137] Sp. nov Valid Bucur & Fucelli in Bucur et al. Late Triassic (Norian)  United States
( Nevada)
A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Holosporella? rossanae[137] Sp. nov Valid Bucur & Del Piero in Bucur et al. Late Triassic (Norian)  Canada
( Yukon)
A green alga belonging to the group Dasycladales and the family Triploporellaceae.

Inocladus[149]

Gen. et comb. nov

Valid

LoDuca et al.

Silurian

 United States

An alga related to the group Bryopsidales. Genus includes "Buthotrephis" divaricata White (1901), "B." newlini White (1901), "B." lesquereuxi Grote & Pitt (1876) and "Chondrites" verus Ruedemann (1925). Announced in 2020; the final version of the article naming it was published in 2021.

Intermurella ordosensis[133]

Sp. nov

Valid

Zhang et al.

Late Ordovician

Ordos Basin

 China

A green alga belonging to the group Dasycladales.

Jurafructus[150]

Gen. et sp. nov

Valid

Chen et al.

MiddleLate Jurassic

Jiulongshan Formation

 China

A plant of uncertain phylogenetic placement, possibly a flowering plant described on the basis of a probable fossil drupe. Genus includes new species J. daohugouensis.

Keraphyton[151]

Gen. et sp. nov

Valid

Champreux, Meyer-Berthaud & Decombeix

Devonian (Famennian)

Mandowa Mudstone Formation

 Australia

A member of Iridopteridales of uncertain phylogenetic placement. Genus includes new species K. mawsoniae.

Khasurtya[152]

Gen. et sp. nov

Valid

Mamontov in Kopylov et al.

Early Cretaceous

 Russia

A liverwort belonging to the group Marchantiidae. Genus includes new species K. ginkgoides.

Lepidodendron demkinae[153]

Sp. nov

Valid

Mosseichik

Carboniferous (Viséan)

 Russia

Lobatannularia linjiaensis[154]

Sp. nov

Valid

Xu et al.

Middle Triassic

Linjia Formation

 China

A member of Equisetales.

Lygodium sanshuiense[155]

Sp. nov

Valid

Naugolnykh et al.

Paleocene

Buxin Formation

 China

A species of Lygodium. Announced in 2019; the final version of the article naming it was published in 2020.

Metzgeriites kujiensis[156]

Sp. nov

Valid

Katagiri in Katagiri & Shinden

Late Cretaceous (Santonian)

Tamagawa Formation

 Japan

A liverwort.

Munieria martinclosasi[134]

comb. nov.

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Lebanon

 Lebanon

Clavatoracean thallus. Formerly known as Charaxis martinclosasi

Neoarthropitys[157] Gen. et sp. nov In press Gnaedinger et al. Middle Triassic Quebrada de los Fósiles  Argentina A member of Equisetales. Genus includes new species N. gondwanaensis.

Osmundacaulis asiatica[158]

Sp. nov

Valid

Cheng et al.

Cretaceous

 China

A member of the family Osmundaceae

Osmundacaulis sinica[158]

Sp. nov

Valid

Cheng et al.

Cretaceous

 China

A member of the family Osmundaceae

Ovoidites circumplicatus[159]

Sp. nov

Valid

Zavattieri, Gutiérrez & Monti

Middle Triassic

Quebrada de los Fósiles Formation

 Argentina

A green alga belonging to the group Zygnematales.

Ovoidites tripartitus[159]

Sp. nov

Valid

Zavattieri, Gutiérrez & Monti

Middle Triassic

Quebrada de los Fósiles Formation

 Argentina

A green alga belonging to the group Zygnematales.

Palaeostachya guanglongii[160]

Sp. nov

In press

Liu et al.

Permian (Asselian)

Taiyuan Formation

 China

A member of the family Calamitaceae.

Parazosterophyllum[146]

Gen. et sp. nov

Valid

McSweeney, Shimeta & Buckeridge

Late Silurian–early Devonian

 Australia

A member of Zosterophyllaceae. Genus includes new species P. timsiae.

Patruliuspora oregonica[137] Sp. nov Valid Bucur & Rigaud in Bucur et al. Late Triassic (Norian)  United States
( Oregon)
A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Patruliuspora pacifica[137] Sp. nov Valid Bucur, Del Piero & Peyrotty in Bucur et al. Late Triassic (Norian)  Canada
( Yukon)
A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Pellites hamiensis[161] Sp. nov Valid Li et al. Middle Jurassic Xishanyao  China A liverwort belonging to the family Pelliaceae.

Plenasium (Aurealcaulis) elegans[162]

Sp. nov

In press

Hiller et al.

Eocene

Na Duong Formation

 Vietnam

A member of Osmundaceae

Pleuromeia shaolinii[163]

Sp. nov

Valid

Zhang & Wang in Zhang et al.

Middle Triassic

Linjia

 China

Polycingulatisporites multiverrucata[136]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla

 Argentina

Spores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version of the article naming it was published in 2020.

Polypodiisporites minutiverrucatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.

Polysporia baetica[164]

Sp. nov

In press

Álvarez-Vázquez, Bek & Drábková

Carboniferous (Pennsylvanian)

Peñarroya-Belmez-Espiel Coalfield

 Spain

A member of Isoetales

Polystichum pacltovae[165]

Sp. nov

Valid

Kvaček in Kvaček & Teodoridis

Oligocene

 Czech Republic

A fern, a species of Polystichum

Proodontosoria[166]

Gen. et sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A fern belonging to the family Lindsaeaceae. Genus includes new species P. myanmarensis.

Proterocladus antiquus[167]

Sp. nov

Valid

Tang et al.

Mesoproterozoic
circa 1 Ga

Nanfen

 China

An early siphonocladalean chlorophyte

Psilatriletes cozzuolii[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.

Psilochara monevaensis[168]

Sp. nov

Valid

Sanjuan & Soulié-Märsche

Middle Miocene

 Spain

A charophyte.

Qianshouia[169]

Gen. et sp. nov

Valid

Huang et al.

Late Devonian

Wutong Formation

 China

A plant of uncertain phylogenetic placement, possibly a lycopsid or a sphenopsid. Genus includes new species Q. mira.

Scolecopteris minuta[170]

Sp. nov

In press

Wan et al.

Early Permian

Taiyuan Formation

 China

A fern belonging to the group Marattiales.

Sigillaria pfefferkornii[171]

Sp. nov

In press

D'Antonio, Boyce & Wang

Permian (Asselian)

 China

Sigillaria wudensis[171]

Sp. nov

In press

D'Antonio, Boyce & Wang

Permian (Asselian)

 China

Sphaerochara miocenica[168]

Sp. nov

Valid

Sanjuan & Soulié-Märsche

Miocene

 Lebanon
 Spain

A charophyte.

Thyrsopteris cretacea[172]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A species of Thyrsopteris

Tomiostrobus sinensis[173]

Sp. nov

Valid

Feng in Feng et al.

Early Triassic (Induan)

Kayitou Formation

 China

A member of the family Isoetaceae.

Tumidopteris astra[174]

Sp. nov

Valid

Naugolnykh

Permian (Roadian)

Pechora coal basin

 Russia

A fern belonging to the family Gleicheniaceae.

Ufadendron elongatum[175]

Sp. nov

Valid

Tang et al.

Late Permian

Linxi

 China

A lycopsid belonging to the family Tomiodendraceae

Uzhurodendron[176]

Gen. et sp. nov

Valid

Mosseichik & Filimonov

Carboniferous (Tournaisian)

Bystrianskaya

 Russia
( Krasnoyarsk Krai)

A member of Lycopodiopsida. Genus includes new species U. asiaticum.

Zeilleria fosteri[177]

Sp. nov

Valid

Thomas et al.

Carboniferous (Bashkirian)

 United Kingdom

A fern

General research[edit]

  • A study on the evolutionary history of green plants is published by Nie et al. (2020).[178]
  • Description of new fossil material of Yurtusia uniformis from the Cambrian Yanjiahe Formation (China) and a study on the phylogenetic relationships and possible life cycle of this organism is published by Shang et al. (2020), who consider Y. uniformis to be a likely green microalga.[179]
  • A study on the phylogenetic relationships of extant and fossil complex thalloid liverworts (Marchantiidae) is published by Flores et al. (2020).[180]
  • Evidence of development of dichotomous roots in euphyllophytes that were extant during the Devonian and Carboniferous periods is presented by Hetherington, Berry & Dolan (2020), who interpret their findings as indicating that dichotomous root branching evolved in both lycophytes and euphyllophytes.[181]
  • An early land plant producing multiple spore size classes is described from the Lower Devonian Campbellton Formation (Canada) by Bonacorsi et al. (2020).[182]
  • A study on the impact of the appearance and evolution of herbivorous tetrapods on the evolution of land plants from the Carboniferous to the Early Triassic is published by Brocklehurst, Kammerer & Benson (2020).[183]
  • A study on the production of periderm in Late Paleozoic arborescent lycopsids is published by D'Antonio & Boyce (2020), who argue that these lycopsids did not grow from sporelings into large trees through the production of a periderm cylinder, because physiological limitations would have prohibited the production of thick periderm.[184]
  • A study on the architecture and development of the Carboniferous arborescent lycopsid Paralycopodites is published by DiMichele & Bateman (2020).[185]
  • New information on the anatomy of Weichselia reticulata is presented by Blanco‐Moreno, Decombeix & Prestianni (2020).[186]
  • A study on the phylogenetic placement of the extinct fern genus Coniopteris is published by Li et al. (2020).[187]
  • New information on the morphology of Paleoazolla patagonica is presented by Benedetti et al. (2020), who evaluate the implications of this taxon for the knowledge of the evolution of water ferns.[188]
  • A study aiming to determine which ferns were most likely to be the producers of Cyathidites spores from earliest Paleocene plant localities across western North America, and were most likely to be among the first plants in western North America to thrive in the immediate aftermath of the Cretaceous–Paleogene extinction event, is published by Berry (2020).[189]
  • A study on the morphology and development of Genomosperma, and on its implications for the knowledge of the evolutionary origins of seed development, is published by Meade, Plackett & Hilton (2020).[190]
  • A pollen organ resembling seed fern pollen organs Dictyothalamus and Melissiotheca is described from the Lopingian Umm Irna Formation (Jordan) by Zavialova et al. (2020), who interpret this finding as evidence of persistence of lyginopterid seed ferns until the late Permian.[191]
  • Evidence of increasing atmospheric CO2 concentration at the onset of the end-Triassic extinction event, based on data from fossil leaves of the seed fern Lepidopteris ottonis from southern Sweden, is presented by Slodownik, Vajda & Steinthorsdottir (2020), who confirm L. ottonis as a valid proxy for pCO2 reconstructions.[192]
  • A study on the anatomy of the seed cone scales of Krassilovia mongolica is published by Herrera et al. (2020), who argue that K. mongolica and Podozamites harrisii are the seed cones and leaves of the same extinct plant, and name a new family Krassiloviaceae within the order Voltziales.[193]
  • A study on the microscopic wood anatomy of a fossil tree trunk of Agathoxylon arizonicum with the characteristic external features of a fire scar from the Upper Triassic Chinle Formation (Petrified Forest National Park, Arizona, United States) is published by Byers et al. (2020), who evaluate the implications of this specimen for the knowledge of the evolution of fire-adapted plant traits.[194]
  • A putative bamboo "Chusquea" oxyphylla from the early Eocene Laguna del Hunco biota (Argentina) is reinterpreted as a conifer by Wilf (2020), who transfers this species to the genus Retrophyllum.[195]
  • A study on evolutionary history of conifers as indicated by fossil and molecular data, aiming to determine whether the rise of angiosperms drove the decline of conifers and other gymnosperms, is published by Condamine et al. (2020).[196]
  • Presence of secretory tissues is reported in extinct flowers from the Cretaceous amber from Myanmar and Cenozoic Dominican amber (including specimens preserved while in the process of emitting compounds) by Poinar & Poinar (2020).[197]
  • Fossil pollen of flowering plants is reported from the Aptian and Albian of Australia by Korasidis & Wagstaff (2020), who evaluate the implications of their findings for the knowledge of the timing of the appearance and diversification of the flowering plants in the high-latitude southern basins of Australia.[198]
  • A study on the morphology of palm and palm-like pollen from the Eocene Yaw Formation (Myanmar), and on the implications of these fossils for the knowledge of distribution and diversity of Eocene palms across the globe, is published by Huang et al. (2020).[199]
  • Fossils fruits of Illigera eocenica, representing the second fossil occurrence of Illigera worldwide and the first in Asia, are described from the Eocene Niubao Formation (central Tibetan Plateau) by Wang et al. (2020), who evaluate the implications of this finding for the knowledge of the climate in the central Tibetan Plateau during the early middle Eocene, and for the knowledge of the floristic links between Asia and North America during the Paleogene.[200]
  • A study on the morphology and phylogenetic relationships of Montsechia vidalii is published by Gomez et al. (2020).[201]
  • Eocene leaves of members of the family Urticaceae with stinging trichomes are described from the Okanogan Highlands (British Columbia, Canada) by DeVore et al. (2020).[202]
  • A revision of the fossil record of the family Nothofagaceae from South America is published by Pujana et al. (2020).[203]
  • A study on the extinction of plants from south polar terrestrial ecosystems during the Permian–Triassic extinction event and on their recovery after this extinction event, based on data from the Sydney Basin (Australia), is published by Mays et al. (2020).[204]
  • A study on the impact of ecological disturbances around the Permian–Triassic boundary (from the Wuchiapingian to Ladinian) on land plant communities is published by Nowak, Vérard & Kustatscher (2020).[205]
  • A study on the age of the Paleogene Kanaka Creek fossil flora (Huntingdon Formation; British Columbia, Canada) and on its implications for reconstructions of the contemporaneous paleoclimate and paleoenvironment is published by Mathewes, Greenwood & Love (2020).[206]
  • Evidence from Eocene plant fossils from the Bangong-Nujiang suture indicating that the Tibetan Plateau area hosted a diverse subtropical ecosystem approximately 47 million years ago and that this area was both low and humid at the time is presented by Su et al. (2020), who also report that the composition of this flora is similar to Early-Middle Eocene floras in both North America and Europe, but shows little affinity to Eocene floras from the Indian Plate.[207]
  • A study aiming to estimate leaf dry mass per area in fossil plants from 22 western North American sites spanning the Eocene–Oligocene transition is published online by Butrim & Royer (2020), who evaluate the implications of their findings for the knowledge of the impact of the environmental changes occurring during the Eocene–Oligocene transition on leaf‐economic strategies of plants.[208]
  • A study on the Neogene paleobotanical record and climate in the northernmost part of the Central Andean Plateau, based on data from the Descanso Formation (Peru), is published by Martínez et al. (2020), who report the earliest evidence of a puna-like ecosystem in the Pliocene and a montane ecosystem without modern analogs in the Miocene.[209]
  • Fossil fruits (mericarps) of the neoendemic Apiaceae Melanoselinum (Daucus) decipiens were reported from the lacustrine and fluvial sediments of Porto da Cruz, Madeira, dated 1.3 Ma, by Góis-Marques et al. 2020.[210] This paper not only reports the oldest Daucus s.l. fossil known to date but also the first fossil evidence of a plant with insular woodiness (see Island gigantism).
  • The leaf fossil Mesodescolea plicata from the Early Cretaceous of Patagonia, first interpreted as a cycad with affinities with extant Stangeria, is reinterpreted as an angiosperm leaf with affinities with Austrobaileyales or Chloranthales by Coiro et al. 2020,[211] with implications for the evolution of leaf shape in the early radiation of the angiosperms.
  • A study on the phylogenetic relationships of 10 Cretaceous flower taxa (Chloranthistemon endressii, Dakotanthus cordiformis, Kajanthus lusitanicus, Mauldinia mirabilis, Microvictoria svitkoana, Paleoclusia chevalieri, Paradinandra suecica, Spanomera mauldiniensis, Tylerianthus crossmanensis and Virginianthus calycanthoides) is published by Schönenberger et al. (2020).[212]

References[edit]

  1. ^ Shook Ling Low; Tao Su; Teresa E. V. Spicer; Fei-Xiang Wu; Tao Deng; Yao-Wu Xing; Zhe-Kun Zhou (2020). "Oligocene Limnobiophyllum (Araceae) from the central Tibetan Plateau and its evolutionary and palaeoenvironmental implications". Journal of Systematic Palaeontology. 18 (5): 415–431. Bibcode:2020JSPal..18..415L. doi:10.1080/14772019.2019.1611673. S2CID 208589882.
  2. ^ a b Yuling Na; Jane Blanchard; Hongshan Wang (2020). "Fruits, seeds and flowers from the Puryear clay pit (middle Eocene Cockfield Formation), western Tennessee, USA". Palaeontologia Electronica. 23 (3): Article number 23(3):a49. doi:10.26879/1045.
  3. ^ a b c d e Vann Smith; Sophie Warny; David M. Jarzen; Thomas Demchuk; Vivi Vajda; Sean P.S. Gulick (2020). "Paleocene–Eocene palynomorphs from the Chicxulub impact crater, Mexico. Part 2: angiosperm pollen". Palynology. 44 (3): 489–519. Bibcode:2020Paly...44..489S. doi:10.1080/01916122.2019.1705417. S2CID 213827225.
  4. ^ Mahasin Ali Khan; Manoshi Hazra; Sumana Mahato; Robert A. Spicer; Kaustav Roy; Taposhi Hazra; Manosij Bandopadhaya; Teresa E.V. Spicer; Subir Bera (2020). "A Cretaceous Gondwana origin of the wax palm subfamily (Ceroxyloideae: Arecaceae) and its paleobiogeographic context". Review of Palaeobotany and Palynology. 283: Article 104318. Bibcode:2020RPaPa.28304318K. doi:10.1016/j.revpalbo.2020.104318. S2CID 224946279.
  5. ^ Mahasin Ali Khan; Kaustav Roy; Taposhi Hazra; Sumana Mahato; Subir Bera (2020). "A new coryphoid palm from the Maastrichtian-Danian sediments of Madhya Pradesh and its palaeoenvironmental implications". Journal of the Geological Society of India. 95 (1): 75–83. Bibcode:2020JGSI...95...75K. doi:10.1007/s12594-020-1388-1. S2CID 210134584.
  6. ^ David Robert Greenwood; John G. Conran (2020). "Fossil coryphoid palms from the Eocene of Vancouver, British Columbia, Canada". International Journal of Plant Sciences. 181 (2): 224–240. doi:10.1086/706450. S2CID 208587364.
  7. ^ Kaustav Roy; Taposhi Hazra; Manoshi Hazra; Sumana Mahato; Subir Bera; Mahasin Ali Khan (2020). "A new coryphoid costapalmate palm leaf from the Maastrichtian-Danian of India". Botany Letters. 168 (2): 155–166. doi:10.1080/23818107.2020.1845974. S2CID 229408918.
  8. ^ Patricia Vallati; Andrea De Sosa Tomas; Gabriel Casal (2020). "A Maastrichtian terrestrial palaeoenvironment close to the K/Pg boundary in the Golfo San Jorge basin, Patagonia, Argentina". Journal of South American Earth Sciences. 97: Article 102401. Bibcode:2020JSAES..9702401V. doi:10.1016/j.jsames.2019.102401.
  9. ^ a b c d e f g h Zlatko Kvaček; Vasilis Teodoridis; Thomas Denk (2020). "The Pliocene flora of Frankfurt am Main, Germany: taxonomy, palaeoenvironments and biogeographic affinities". Palaeobiodiversity and Palaeoenvironments. 100 (3): 647–703. Bibcode:2020PdPe..100..647K. doi:10.1007/s12549-019-00391-6.
  10. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj Fátima Praxedes Rabelo Leite; Silane Aparecida Ferreira da Silva-Caminha; Carlos D’Apolito (2020). "New Neogene index pollen and spore taxa from the Solimões Basin (Western Amazonia), Brazil". Palynology. 45 (1): 115–141. doi:10.1080/01916122.2020.1758971. S2CID 219090032.
  11. ^ M. Laura Pipo; Ari Iglesias; Josefina Bodnar (2020). "A new vesselless angiosperm stem with a cambial variant from the Upper Cretaceous of Antarctica". Acta Palaeontologica Polonica. 65 (2): 261–272. doi:10.4202/app.00697.2019. hdl:11336/136234.
  12. ^ a b c d Indah B. Huegele; Steven R. Manchester (2020). "An early Paleocene carpoflora from the Denver Basin of Colorado, USA, and its implications for plant-animal interactions and fruit size evolution". International Journal of Plant Sciences. 181 (6): 646–665. doi:10.1086/707727. S2CID 219748982.
  13. ^ a b c d Lilla Hably (2020). "The Karpatian (late early Miocene) flora of the Mecsek area". Acta Palaeobotanica. 60 (1): 51–122. doi:10.35535/acpa-2020-0003.
  14. ^ Thomas Denk; Johannes Martin Bouchal; Pavel Smirnov; Yaroslav Trubin (2020). "Late Oligocene leaf and pollen flora of Southwestern Siberia: taxonomy, biogeography and palaeoenvironments". Historical Biology: An International Journal of Paleobiology. 33 (11): 2951–2976. doi:10.1080/08912963.2020.1839064. S2CID 230547821.
  15. ^ Hai Zhu; Steven R. Manchester (2020). "Fruit of Staphylea (Staphyleaceae) from the Oligocene of Montana, USA". Review of Palaeobotany and Palynology. 280: Article 104275. Bibcode:2020RPaPa.28004275Z. doi:10.1016/j.revpalbo.2020.104275. S2CID 225729365.
  16. ^ Susanne S. Renner; Viviana D. Barreda; María Cristina Tellería; Luis Palazzesi; Tanja M. Schuster (2020). "Early evolution of Coriariaceae (Cucurbitales) in light of a new early Campanian (ca. 82 Mya) pollen record from Antarctica". Taxon. 69 (1): 87–99. doi:10.1002/tax.12203.
  17. ^ MacKenzie Allan Smith; Steven R. Manchester (2020). "CT-scans of capsules from the Clarno Formation (Oregon, USA) reveal an extinct Eocene theaceous taxon". Acta Palaeobotanica. 60 (2): 251–258. doi:10.35535/acpa-2020-0013.
  18. ^ Xiang‐Chuan Li; Steven R. Manchester; Qin Wang; Liang Xiao; Tian‐Long Qi; Yun‐Zhi Yao; Dong Ren; Qiang Yang (2020). "A unique record of Cercis from the late early Miocene of interior Asia and its significance for paleoenvironment and paleophytogeography". Journal of Systematics and Evolution. 59 (6): 1321–1338. doi:10.1111/jse.12640. S2CID 219523602.
  19. ^ a b Johanna Baez; Alexandra Crisafulli (2020). "Novelties in the xylotaphoflora from Chiquimil Formation (Miocene), Catamarca-Argentina". Journal of South American Earth Sciences. 107: Article 102943. doi:10.1016/j.jsames.2020.102943. S2CID 225109309.
  20. ^ a b Oris J. Rodríguez-Reyes; Emilio Estrada-Ruiz (2020). "Two new reports of ancient rainforest trees from the Azuero Peninsula, Panama". Ameghiniana. 57 (3): 209–218. doi:10.5710/AMGH.22.02.2020.3299. S2CID 216250364.
  21. ^ a b c A.L. Averyanova; Yaowu Xing (2020). "New Paleogene angiosperm species of Zaissan Depression (eastern Kazakhstan)". Botanicheskii Zhurnal. 105 (1): 46–57. doi:10.31857/S0006813620010044. S2CID 218793001.
  22. ^ a b c d e f g h Xiaoyan Liu; Hanzhang Song; Jianhua Jin (2020). "Diversity of Fagaceae on Hainan Island of South China during the middle Eocene: implications for phytogeography and paleoecology". Frontiers in Ecology and Evolution. 8: Article 255. doi:10.3389/fevo.2020.00255. S2CID 221117336.
  23. ^ a b Li Xue; Linbo Jia; Gi-soo Nam; Yongjiang Huang; Shitao Zhang; Yuqing Wang; Zhuo Zhou; Yongsheng Chen (2020). "Involucre fossils of Carpinus, a northern temperate element, from the Miocene of China and the evolution of its species diversity in East Asia". Plant Diversity. 42 (3): 155–167. doi:10.1016/j.pld.2020.01.001. PMC 7361179. PMID 32695948.
  24. ^ Anthony L. Swinehart; James O. Farlow (2020). "Plant and invertebrate macrofossils from the Pipe Creek Sinkhole (Late Neogene), Grant County, Indiana". Historical Biology: An International Journal of Paleobiology. 33 (11): 3111–3140. doi:10.1080/08912963.2020.1851686. S2CID 230538832.
  25. ^ Dimitra Mantzouka; Miroslav Ivanov; Vladimir Bozukov (2020). "The first report of an 'evergreen Castanopsis type' wood (Fagaceae) for the Late Miocene–Early Pliocene of Europe (Bulgaria, Blagoevgrad Graben)". Geological Journal. 56 (2): 786–803. doi:10.1002/gj.3919.
  26. ^ Cédric Del Rio; Teng‐Xiang Wang; Jia Liu; Shui‐Qing Liang; Robert A. Spicer; Fei‐Xiang Wu; Zhe‐Kun Zhou; Tao Su (2020). "Asclepiadospermum gen. nov., the earliest fossil record of Asclepiadoideae (Apocynaceae) from the early Eocene of central Qinghai‐Tibetan Plateau, and its biogeographic implications". American Journal of Botany. 107 (1): 126–138. doi:10.1002/ajb2.1418. PMID 31944266.
  27. ^ Cédric Del Rio; Jian Huang; Gregory W. Stull; Rémi Allemand; Zhe‐Kun Zhou; Tao Su (2020). "First macrofossil record of Icacinaceae in East Asia (early Oligocene, Wenshan Basin) and its ecological implications". Journal of Systematics and Evolution. 60 (2): 445–455. doi:10.1111/jse.12700. S2CID 228976920.
  28. ^ Cédric Del Rio; Dario De Franceschi (2020). "Icacinaceae fossil fruits from three sites of the Paris Basin (early Eocene, France): local diversity and global biogeographic implications". Geodiversitas. 42 (2): 17–28. doi:10.5252/geodiversitas2020v42a2.[permanent dead link]
  29. ^ Andrew C. Rozefelds; Gregory Stull; Peta Hayes; David R. Greenwood (2020). "The fossil record of Icacinaceae in Australia supports long-standing Palaeo-Antarctic rainforest connections in southern high latitudes". Historical Biology: An International Journal of Paleobiology. 33 (11): 2854–2864. doi:10.1080/08912963.2020.1832089. S2CID 229005088.
  30. ^ Gregory Stull; Bruce H. Tiffney; Steven R. Manchester; Cédric Del Rio; Scott L. Wing (2020). "Endocarps of Pyrenacantha (Icacinaceae) from the early Oligocene of Egypt". International Journal of Plant Sciences. 181 (4): 432–442. doi:10.1086/706854. S2CID 208558190.
  31. ^ Ünal Akkemik; Dimitra Mantzouka; Umut Tunç; Fikret Koçbulut (2020). "The first paleoxylotomical evidence from the Mid-Eocene Climate Optimum from Turkey". Review of Palaeobotany and Palynology. 285: Article 104356. doi:10.1016/j.revpalbo.2020.104356. S2CID 229395041.
  32. ^ a b c Daniela P. Ruiz; M. Sol Raigemborn; Mariana Brea; Roberto R. Pujana (2020). "Paleocene Las Violetas Fossil Forest: Wood anatomy and paleoclimatology". Journal of South American Earth Sciences. 98: Article 102414. Bibcode:2020JSAES..9802414R. doi:10.1016/j.jsames.2019.102414. S2CID 213796947.
  33. ^ Kenton L. Chambers; George O. Poinar, Jr (2020). "Thymolepis toxandra gen. et sp. nov., a mid-Cretaceous fossil flower with horseshoe-shaped anthers". Journal of the Botanical Research Institute of Texas. 14 (1): 57–64. doi:10.17348/jbrit.v14.i1.896.
  34. ^ George O. Poinar, Jr; Kenton L. Chambers; Urszula T. Iwaniec; Fernando E. Vega (2020). "Valviloculus pleristaminis gen. et sp. nov., a Lauralean fossil flower with valvate anthers from mid-Cretaceous Myanmar amber". Journal of the Botanical Research Institute of Texas. 14 (2): 359–366. doi:10.17348/jbrit.v14.i2.1014.
  35. ^ a b Jun‐Ling Dong; Bai‐Nian Sun; Ai‐Jing Li; Hui Chen (2020). "The diversity of Smilax (Smilacaceae) leaves from the Middle Miocene in southeastern China". Geological Journal. 56 (2): 744–757. doi:10.1002/gj.3882. S2CID 225790148.
  36. ^ Lu‐Liang Huang; Jian‐Hua Jin; Cheng Quan; Alexei A. Oskolski (2020). "Mummified Magnoliaceae woods from the upper Oligocene of South China, with biogeography, paleoecology, and wood trait evolution implications". Journal of Systematics and Evolution. 58 (1): 89–100. doi:10.1111/jse.12480. S2CID 91861708.
  37. ^ Else Marie Friis; Peter R. Crane; Kaj Raunsgaard Pedersen (2020). "Melloniflora, a new extinct multiparted flower from the Early Cretaceous of Virginia, USA". International Journal of Plant Sciences. 181 (9): 887–897.